Carol Berman


Carol Berman.

Carol Berman


Carol Berman


Research Areas

Primate social behavior; Parent-offspring relationships; Animal behavior; Evolution of behavior; Kinship; Conflict management; Ethological methods.


  • PhD, Cambridge University, 1978  

Recent Publications

A.    Book
B. Chapais and C. M. Berman.  KINSHIP AND BEHAVIOR IN PRIMATES. Oxford University Press, New York, 520 pp., 2004.

B. Peer Reviewed Publications (full length journal articles and chapters)

L.R. LaBarge, A. Alan, C.M. Berman, R. Hill and S. Margulis. Anthropogenic influences on primate antipredator behavior and implications for research and conservation. AMERICAN JOURNAL OF PRIMATOLOGY, in press.

K. Balasubramaniam, H. Ogawa, J-H Li and C.M. Berman. Tibetan Macaque Social Style: Co-variant and Quasi-independent Evolution. In: The Behavioral Ecology of Tibetan Macaques, ed. by L. Sun, J-H. Li and P. Garber, Springer Press. Pp. 141-171, 2020.

E. Dunayer & C.M. Berman. Time matching between grooming partners: Do distinctions between pauses vs. bout terminations matter? AMERICAN JOURNAL OF PRIMATOLOGY, online DOI: 10.1002/ajp.22968, 2019
E. Dunayer & C.M. Berman. Infant-Directed Handling Among Primates. INTERNATIONAL JOURNAL OF COMPARATIVE PSYCHOLOGY, Special Issue, Vol. 31. 2018.

A. Yanagi & C.M. Berman. Nonhuman primate social play: Coping with costs. In The Cambridge Handbook of Play: Developmental and Disciplinary Perspectives. Edited by P. K. Smith and J.L.Roopnarine, Cambridge University Press, pp. 67-83, 2018. DOI:10.1017/9781108131384.

K.N. Balasubramaniam, B.A. Beisner, C.M. Berman, A. De Marco, J. Duboscq, S. Koirala,  B. Majolo, A.J.J. MacIntosh, R. McFarland, S. Molesti,  H.Ogawa, O. Petit, G. Schino, S. Sosa, B. Thierry, F.B.M. de Waal, & B.J. McCowan. The influence of phylogeny, social style and sociodemographic factors on macaque social networks. AMERICAN JOURNAL OF PRIMATOLOGY, 80: e22727, 2018. DOI: 10.1002/ajp.22727, 2017.

K.N. Balasubramaniam & C.M. Berman. Market-based Grooming Exchange for Resource Tolerance in a free-ranging Group of Rhesus Macaques at Cayo Santiago, BEHAVIOUR, 154: 1145-1176, 2017. DOI:10.1163/1568539X-00003462.

E.A. Weigel & C.M. Berman. Body signals used during social play in captive immature western lowland gorillas (Gorilla gorilla gorilla).  PRIMATES, 59: 253-265, 2018.
E. Dunayer & C.M. Berman. Infant handling enhances social bonds in free-ranging rhesus macaques (Macaca mulatta). BEHAVIOUR, 154: 875-907, 2017. DOI:10.1163/1568539X-00003448.

A. Yanagi & C.M. Berman. Does behavioural flexibility contribute to successful play
among juvenile rhesus macaques? BEHAVIOURAL ECOLOGY AND SOCIOBIOLOGY, 71: 156-175, 2017. DOI 10.1007/s00265-017-2377-2.

E.S. Dunayer & C.M. Berman.   Theoretical origins and interpretive debates, and proximate perspective: A response to Sánchez-Amaro and Amici. ANIMAL BEHAVIOUR, published online October 3, 2016, printed version: 121: 131-136, 2016.

C.M. Berman. Primate kinship: contributions from Cayo Santiago. AMER. JOURNAL OF PRIMATOLOGY. Invited ms and featured article for a special edition, published online: early view-DOI=10.1002/ajp.22383&ArticleID=2177526, 2015. Special edition—78:63-77, 2016.

K.N. Balasubramaniam , E.S. Dunayer, L.J. Gilhooly, K.A. Rosenfield & C.M. Berman. Group Size, Contest Competition, and Social Structure in Cayo Santiago Rhesus Macaques. BEHAVIOUR, 151: 1759-1798, 2014.

A. Yanagi & C.M. Berman. Functions of multiple play signals in free-ranging juvenile rhesus macaques (Macaca mulatta). BEHAVIOUR, 151: 1983-2014, 2014.

K.N Balasubramaniam, K, Dittmar, C.M. Berman, M., Butovskaya, M.A. Cooper, B. Majolo, H. Ogawa, G. Schino, B.Thierry & Waal. Hierarchical Steepness and Phylogenetic Models I: Phylogenetic Signals in Macaca, ANIMAL BEHAVIOUR., 83: 1207-1218, 2012.

K.N Balasubramaniam, K, Dittmar, C.M. Berman, M., Butovskaya, M.A. Cooper,B. Majolo, H. Ogawa, G. Schino, B.Thierry & Waal. Hierarchical Steepness, Counter-aggression and the Macaque Social Scale. AMERICAN J. PRIMATOLOGY, 74: 915-925, 2012.

 A. Yanagi & C.M. Berman. Gestural signals during social play in free-ranging rhesus macaques (Macaca mulatta): A systematic analysis. AMERICAN JOURNAL OF PRIMATOLOGY, Published online: DOI: 10.1002/ajp.22219, 9/30/13.